What does some symbol in regular express mean?

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x = [1 0.3 -2 0.001 -0.00016, 582398, 3020];
regexp(num2str(x), '(?<=(^|\s+)[\-\.0]*)[1-9](?=\d*\.?\d*)', 'match')
What does these symbol '<' '=' mean?
How this expression work, first do what, then do what?

采纳的回答

Rolfe Dlugy-Hegwer
(?<=expr) - Look behind from current position and test if expr is found.

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