Hi Jacopo,
this SBML file is indeed not valid according to the SBML validator. One unit definition is missing the 'id' attribute.
That said, a valid unit (with an 'id') with the same name and same definition is defined a few lines above it.
So you can safely delete lines 80-85 of the file and save it. The model will then get imported just fine.
This model seems to have a couple of issues though. For example parameter k1_74 does have any unit. Also, the Peripheral and Lymph Nodes compartments are not used and do not contain any species. But there might be more information about this in the associated paper.
Best regards,
Jérémy